They discovered that no regular cerevisiae or carlsbergensis strains utilised lysine whereas many other yeasts, together with wild yeasts, did so. They stored their stock cultures on malt extract agar slopes or on malt extract chalk agar in the case of Brettanomyces species. Later, Morris & Eddy1 described a solid lysine medium for the isolation and enumeration of wild yeasts in pitching yeast.

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